It's so easy to lose sight of the trees in the forest.
The switch is nothing but a conduit between the router and the network host - since you cannot configure the router for jumbo frame, when the router and the client negotiate the MSS it will be 1500 bits so that will be the size of the packet being sent by the host, and so the router will not fragment it.
Let me see if I can make this a little clearer.
Imagine if you will, that you have a router and a five port switch, two separate devices - you plug the router into port 5 on the switch and you plug the client system into port 1 on the switch.
The client system allows you to configure a jumbo frame with a size of 4000 bits, the router does not allow jumbo frame - the two devices automatically negotiate the MSS, which will be 1500 bits (because the router cannot accomodate a larger frame), so the client system will send 1500 bit frames, and the router will not fragment them.
Let us now connect the DNS-323 to the five port switch, let's plug that in to port 2 - the client system and the DNS-323 will now negotiate the MSS, which will again be 1500 bits and the two will communicate with no trouble - the user opens a browser on the client system and logs into the DNS-323 and configures that for jumbo frame (4000 bits) and now the two will negotiate an MSS with a 4000 bit limit.
Let us not forget that it is entirely possible to have devices with different frame sizes on the same LAN - if it weren't there would be no way to configure the DNS-323 for jumbo frame (either one device or the other HAS to be set for jumbo frame first)
Get the idea?
OK - now let's squash our router and our five port switch into one cabinet - presto we have a DGL-4500.