My wife uses Norton360 on her machine and her backups are written the our DNS-323. I have taken a quick look at the structure of the output files and is is my guess that the data is stored in a proprietary database. I believe the backups are incremental and it would be my second guess that when a file changes, the old data records are flagged as outdated, the new data is written in a new location and pointers to the data are adjusted. Therefore, after two years your backup file probably has many unused sectors.
An approach to see if this is true would be to delete the N360_BACKUP directory, and recreate a new backup to see how your space utilization changes. Since you still have the files on the Windows machine, you should be fairly safe. It will probably take about 4 or 5 hours to recreate the backup.
Please tell us what you learn.