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I would argue it has more to do with the controllers involved, but that's just my opinion. It's still pretty week to get more performance out of an USB external drive than an Gigabit NAS.
The best place to put your money to optimize NAS performance is to beef up the RAM on a NAS and each networked client.
I reset the DNS-323 to factory defaults. Gigabit speeds went up to 24MB/s again!! ...
Also, what size do you recommend for the Jumbo frames? I've read recommendations on this forum to set it at 4000, what if I max it out?