My unit says the box is at 45C and the drives are at 35C, taken when the box is at idle. That accounts for mine being quiet, both temps fall under the "off" variable when they aren't being accessed. The temps are also taken in a room that is a consistent 22C in the summertime and around 20C in the winter. It is also taken with the stock front cover, no mods, and the cover is fully closed at all times.
As for where the temps are taken, that is a good question. I would have placed a sensor at the exhaust port for the box temp for my preference. But I'm guessing that the 321 has it on the circuit board and the drives have their own internal sensor to give the S.M.A.R.T. info a place to get the temp from. Just a guess though about the box sensor, I'm not sure where Dlink placed it?
Ryder